Turtles are reptiles of the order Chelonii or Testudines characterized by a special bony or cartilaginous shell developed from their ribs and acting as a shield. The earliest known turtles date from 220 million years ago, making turtles one of the oldest reptile groups and a more ancient group than lizards, snakes or crocodiles. Of the many species alive today, some are highly endangered. Turtles are ectotherms—their internal temperature varies according to the ambient environment, commonly called cold-blooded. However, because of their high metabolic rate, leatherback sea turtles have a body temperature that is noticeably higher than that of the surrounding water. Turtles are classified as amniotes, along with other reptiles (including birds) and mammals. Like other amniotes, turtles breathe air and do not lay eggs underwater, although many species live in or around water.