NATURE producer Jill Clarke gives an update on the leadership shakeup in Titus's gorilla family.

While his prolific mating is a clear sign of the king's success, this unique gorilla's extensive family tree grew out of tragic circumstances.

NATURE retraces the life of Titus, silverback leader, from birth to present reign, spanning a tumultuous period in the history of a species and a nation in "The Gorilla King."

Recent challenges to Titus's rule by his rival, Kuryama, have the entire group of gorillas on edge.